This division carries out various aspects of research and education in basic biology such as taxonomy, evolutionary biology, developmental biology, physiological ecology and biogeography from the molecular level to the ecosystem level using mainly temperate marine invertebrates as materials. Moreover, through extension of research fields into the global scale from tropical to boreal regions, the division also covers biology that has applied aspects such as environmental and conservation biology as subjects of research and education.


The laboratory addresses the question of how diverse marine organisms are, and evolve throughout the time, from a view point of natural history sciences. This includes comparative morphology, development, systematics, phylogeny, biogeography, and phylogeography of marine organisms by both morphological and molecular level methods. The laboratory also carries out research on conservation biology of marine organisms of various taxonomic groups from a view point of environmental change and the biotic responses. Seto Marine Biological Laboratory provides this program to undergraduate students in Faculty of Science and students in Marine Biology Course, the Graduate School of Science.

ASAKURA, Akira, D. Sc. (Kyushu Univ.), Invertebrate Zoology; Systematics, ecology, behavior, and biogeography of Crustacea
E-mail : asakura.akira.6w_at_kyoto-u.ac.jp

* Asakura, A. (ed.) “New Frontiers in Crustacean Biology.” Brill, Leiden, The Netherlands, 2011, 351 pp.
* Asakura, A. “Evolution of mating system in decapod crustaceans. In: Joel W. Martin, Keith A. Crandall, and Darryl L. Felder (eds.), Decapod Crustacean Phylogenetics” Taylor & Francis, New York, U.S.A. 2009. p.121-182,
* Asakura, A. “A revision of the hermit crabs of the genera Catapagurus A. Milne-Edwards and Hemipagurus Smith from the Indo-West Pacific (Crustacea: Decapoda: Anomura: Paguridae). p.121-182, Invertebrate Taxonomy. 2001, 15, p.823-891.

■Associate Professor
SHIMOMURA, Michitaka, D. Sc. (Hokkaido Univ.), Invertebrate Zoology; Taxonomy
E-mail : shimomura.michitaka.5w_at_kyoto-u.ac.jp

* Shimomura, M. (2017) A new species of Aspidophryxus (Isopoda, Dajidae), ectoparasitic on Mysidella hoshinoi (Mysidae) in Japan. ZooKeys, 646, 109-118.
* Shimomura, M. (2009) Ten new species of paramunnid isopods (Peracarida: Asellota: Paramunnidae) from Kyushu, southern Japan. Bulletin of the National Museum of Nature and Science, Series A (Zoology), Supplement, 3: 47-88.
* Shimomura, M., Ohtsuka, S. and Naito, K. (2005) Prodajus curviabdominalis n. sp. (Isopoda: Epicaridea: Dajidae), an ectoparasite of mysids, with notes on morphological changes, behavior and life-cycle. Systematic Parasitology, 60: 39-57.

■Assistant Professor
YAMATO, Shigeyuki, D. Sc. (Hiroshima Univ.), Invertebrate Zoology; Taxonomy, Crustacea, Natural History
E-mail : yamato.shigeyuki.5s_at_kyoto-u.ac.jp

* Yusa, Y. ; Yamato, S. ; Marumura, M. (2001) “Ecology of a parasitic barnacle, Koleolepas avis: relationship to the hosts, distribution, left-right asymmetry and reproduction.” J. Mar. Biol. Ass. U.K., 8, p.781-788
* Yamato, S. (1992) “A new species of Podocerus (Amphipoda: Podoceridae) from the carapace of a loggerhead sea turtle in Japan. Publ.” Seto Mar. Biol. Lab., 35(4/5), p.281-288
* Yamato, S. (1987) “Four intertidal species of the genus Melita (Crustacea: Amphipoda) from Japanese waters, including descriptions of two new species. Publ.” Seto Mar. Biol. Lab., 32(4/6), p.275-302

■Assistant Professor
NAKANO, Tomoyuki, D. Sc. (Nagoya Univ.), Invertebrate Zoology; Molecular Phylogeny and Phylogeography
E-mail : nakano.tomoyuki.2a _at_ kyoto-u.ac.jp

* Nakano, T. ; Sasaki, T. ; Kase, T. (2010) “Color polymorphism and historical biogeography in the Japanese patellogastropod limpet Cellana nigrolineata (Reeve) (Patellogastropoda: Nacellidae).” Zoological Science., 27, p.811-820.
* Nakano, T. ; Ozawa, T. (2007) “Worldwide phylogeography of limpets of the order Patellogastropoda: molecular, morphological and palaeontological evidence.” Journal of Molluscan Studies., 73, p.79-99.
* Nakano, T. ; Spencer, H.G. (2007) “Simultaneous polyphenism and cryptic species in an intertidal limpet from New Zealand.” Molecular Phylogenetics and Evolution., 45, p.470-479.

■Assistant Professor
GOTO, Ryutaro, Ph. D in Human Environmental Science (Kyoto Univ.), Evolutionary Biology, Ecology, Marine Biology, Invertebrate Systematics, Symbiosis
E-mail : goto.ryutaro.8n _at_ kyoto-u.ac.jp

* Goto R (2016) A comprehensive molecuar phylogeny of spoon worms (Echiura, Annelida): Implications for morphological evolution, the origin of dwarf males, and habitat shifts. Molecular Phylogenetics and Evolution 99: 247–260.
* Goto R, Kawakita A, Ishikawa H, Hamamura Y, Kato M (2012) Molecular phylogeny of the bivalve superfamily Galeommatoidea (Heterodonta, Veneroida) reveals dynamic evolution of symbiotic lifestyle and interphylum host switching. BMC Evolutionary Biology 12:172.
* Goto R, Kato M (2012) Geographic mosaic of mutually exclusive dominance of obligate commensals in symbiotic communities associated with a burrowing echiuran worm. Marine Biology 159: 319–330


We study to clarify the aquatic ecosystem that maintains the sustainable fisheries production, the mechanism of the change of the ecosystem and the behavior ecology of fishes and aquatic mammals in the aquatic biosphere including marine and fresh waters. In order to research these topics, we perform the stable isotope analysis, the simulation using ecological models and bio-logging techniques including biotelemetry and micro-data loggers. This laboratory is a temporary cooperative laboratory from the Graduate School of Agriculture.

ARAI, Nobuaki, D. Agr. (Kyoto Univ.), Aquatic Biosphere Informatics
E-mail : arai.nobuaki.8c_at_kyoto-u.ac.jp

* Mitamura H. ; Uchida K. ; Miyamoto Y. ; Kakihara T. ; Miyagi A. ; Kawabata Y. ; Ichikawa K. ; Arai N. (2012) “Short-range homing in a site-specific fish: search and directed movements”, J. Exp. Biol., 215, p.2751-2759.
* Noda T. ; Okuyama J. ; Koizumi T. ; Arai N. ; Kobayashi M. (2012) “Monitoring attitude and dynamic acceleration of free-moving aquatic animals using a gyroscope”. Aquat. Biol., 16, p.265-276.
* Okuyama J. ; Kataoka K. ; Kobayashi M. ; Abe O. ; Yoseda K. ; Arai N. (2012) “The regularity of dive performance in sea turtles: a new perspective from precise activity data”. Animal Behaviour., 84, p.349-359.

■Associate Professor
ICHIKAWA, Kotaro, D.Info. (Kyoto Univ.), Underwater bioacoustics and acoustic telemetry
E-mail : ichikawa.kotaro.5r_at_kyoto-u.ac.jp

* Ando-Mizobata, N. ; Ichikawa, K. ; Arai, N. ; Kato, H. (2014) “Does boat noise affect dugong (Dugong dugon) vocalization?” Mammal Study., 36, p.121-127.
* Matsuo, Y. ; Ichikawa, K. ; Ando-Mizobata, N. ; Arai, N. (2013) “Cyclic change of dugong’s vocal behavior.” Journal of Advanced Marine Science and Technology., 19, p.1-4.
* Ichikawa, K. ; Akamatsu, T. ; Arai, N. ; Shinke, T. ; Adulyanukosol, K. (2011) “Callback response of dugongs to conspecific chirp playbacks.” Journal of the Acoustical Society of America., 129, p.3623-3629.

■Assistant Professor
KOBAYASHI, Shiho, D. Agr. (Kyoto Univ.), Marine Science, physical-biological interaction in coastal seas
E-mail : shihok_at_kais.kyoto-u.ac.jp

* Kobayashi, Shiho ; Fujiwara, Tateki ; Zenitani, Hiromu ; Nagamoto, Kazuhisa ; Futamura, Akira. (2010) “Gravitational circulation and its response to the variation in river discharge in the Seto Inland Sea, Japan.” Journal of Geophysical Research., VOL. 115, C03009, doi:10.1029/2007JC004503.
* Kobayashi, Shiho ; Hashimoto, Eisuke ; Nagao, Masayuki ; Takasugi, Yoshio. (2009) “The variation of turbulent energy dissipation and water column stratification at the entrance of a tidally energetic strait.” Journal of Oceanography., 65(5), p.657-664.
* Kobayashi, Shiho ; Fujiwara, Tateki . (2009) “Modeling the long-term variability of shelf water intrusion into the Seto Inland Sea, Japan.” Journal of Marine Research., 77, p.341-349.